After discovering their parents are still alive and their homeland of Gerander is in danger, mouse triplets Alistair, Alice, and Alex, and their friend Tibby Rose, have joined the underground rebel organization FIG. In quick measure, FIG orders Alex and Alice go undercover in Souris to infiltrate Queen Eugenia's palace while Alistair and Tibby Rose are sent to discover Gerander's secret paths, which may be the key Gerandans need to triumph and for the triplets to rescue their parents.

Enemy spies, attacking eagles, and blizzarding mountaintops seem all the more challenging when there is a lack of good cheese available, but these four young mice respond with endless creativity and determination. Cheeky and entertaining, The Spies of Gerander is an action filled sequel to the first book in The Song of the Winns series, The Secret of the Ginger Mice.